A revision of Aceratherium blanfordi Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key
Identifieur interne : 000134 ( France/Analysis ); précédent : 000133; suivant : 000135A revision of Aceratherium blanfordi Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key
Auteurs : Pierre-Olivier Antoine [France] ; Kevin F. Downing [États-Unis] ; Jean-Yves Crochet [France] ; Francis Duranthon [France] ; Lawrence J. Flynn [États-Unis] ; Laurent Marivaux [France] ; Grégoire Métais [France] ; Abdul Rahim Rajpar [Pakistan] ; Ghazala Roohi [Pakistan]Source :
- Zoological Journal of the Linnean Society [ 0024-4082 ] ; 2010-09.
English descriptors
- KwdEn :
- Aceratheriini, Aceratherium, Albigense, Alicornops, Antecrochet, Anterior, Anterior border, Anterior side, Anterior view, Anteroposterior, Anteroposterior diameter, Antoine, Antoine welcomme, Apophysis, Approximate dimensions, Articular, Articulation, Astragalus, Attened, Balochistan, Bicornis, Blanfordi, Bonis, Brachypotherium, Brachypus, Bracket, Browni, Bugti, Bugti hills, Bula, Calcaneum, Calcaneus, Carpalis, Cheek teeth, Chilotherium, Chitarwata, Cingulum, Clade, Coeval, Constricted, Convex, Corpus mandibulae, Cranial, Crista, Cuboid, Deciduous, Dentition, Dera, Dera bugti, Diaceratherium, Diaphysis, Diceratherium, Dicerorhinus, Diceros, Dihedron, Distal articulation, Distal border, Distal view, Dorsal, Dorsal view, Douvillei, Duvernoy, Earliest miocene, Early miocene, Ectocuneiform, Ectoloph, Ectometaloph, Elasmotheriina, Elasmotheriinae, Entoconid, Extensor carpalis, External groove, Facet, Fauna, Femur, Foramen, Fossil, Gaimersheimense, Gaindatherium, Gand, Gers, Ginsburg, Groove, Gutter, Heissig, Homoplastic, Hooijer, Humerus, Hypocone, Hypolophid, Incisivum, Insertion, Interossei, Kumbi, Labial, Labial cingulum, Lartet, Lateral, Lateral side, Lateral view, Laterally, Lholi, Lingual, Lingual bridge, Lingual cingulum, Lingual groove, Linnaeus, Linnean, Linnean society, Lower cheek teeth, Lower molars, Lower premolars, Lydekker, Mammal, Mammalia, Mandible, Mandibulae, Mandibular, Maxilla, Mcii, Mciii, Medial, Medial side, Median, Medifossette, Mesaceratherium, Mesaceratherium welcommi, Mesocuneiform, Mesostyle, Metacarpal, Metacone, Metaconid, Metaloph, Metapodials, Mhnt, Minutum, Miocene, Miocene rhinocerotids, Mirallesi, Mnhn, Molar, Monophyletic, Mtii, Mtiii, Mtiv, Natural history, Naturelle, Navicular, Node, Oblique, Occlusal, Occlusal view, Oligocene, Osborni, Outgroup, Paracone, Paralophid, Patella, Paulhiacense, Perissodactyla, Phylogenetic, Plesiaceratherium, Pleuroceros, Pleuroceros blanfordi, Pmnh, Postcranial, Postcranials, Posterior, Posterior border, Posterior part, Posterior tuberosity, Posterior valley, Posterolateral, Posteroproximal, Praecursor, Premolar, Processus, Processus zygomaticus, Prosantorhinus, Protaceratherium, Protocone, Protoloph, Prox, Proximal, Proximal border, Proximal view, Reversion, Rhinocerotid, Rhinocerotidae, Rhinocerotids, Rhinocerotina, Rhinocerotinae, Right astragalus, Right cuboid, Right scaphoid, Robust, Ronzotherium, Sagittal, Sagittally, Scaphoid, Semilunate, Sensu, Simorrense, Squamosal, Square brackets, Stratigraphical, Suprageneric, Symphysis, Synapomorphies, Talus, Taxon, Tibia, Transverse diameter, Trapezoid, Trigonias, Trigonid, Trochlea, Tuber, Tubercle, Tuberosity, Ulna, Unciform, Upper member, Upper molars, Upper premolars, Welcomme, Welcommi, Zinda, Zoological, Zoological journal, Zygomatic, Zygomatic arch, Zygomaticus.
- Teeft :
- Aceratheriini, Aceratherium, Albigense, Alicornops, Antecrochet, Anterior, Anterior border, Anterior side, Anterior view, Anteroposterior, Anteroposterior diameter, Antoine, Antoine welcomme, Apophysis, Approximate dimensions, Articular, Articulation, Astragalus, Attened, Balochistan, Bicornis, Blanfordi, Bonis, Brachypotherium, Brachypus, Bracket, Browni, Bugti, Bugti hills, Bula, Calcaneum, Calcaneus, Carpalis, Cheek teeth, Chilotherium, Chitarwata, Cingulum, Clade, Coeval, Constricted, Convex, Corpus mandibulae, Cranial, Crista, Cuboid, Deciduous, Dentition, Dera, Dera bugti, Diaceratherium, Diaphysis, Diceratherium, Dicerorhinus, Diceros, Dihedron, Distal articulation, Distal border, Distal view, Dorsal, Dorsal view, Douvillei, Duvernoy, Earliest miocene, Early miocene, Ectocuneiform, Ectoloph, Ectometaloph, Elasmotheriina, Elasmotheriinae, Entoconid, Extensor carpalis, External groove, Facet, Fauna, Femur, Foramen, Fossil, Gaimersheimense, Gaindatherium, Gand, Gers, Ginsburg, Groove, Gutter, Heissig, Homoplastic, Hooijer, Humerus, Hypocone, Hypolophid, Incisivum, Insertion, Interossei, Kumbi, Labial, Labial cingulum, Lartet, Lateral, Lateral side, Lateral view, Laterally, Lholi, Lingual, Lingual bridge, Lingual cingulum, Lingual groove, Linnaeus, Linnean, Linnean society, Lower cheek teeth, Lower molars, Lower premolars, Lydekker, Mammal, Mammalia, Mandible, Mandibulae, Mandibular, Maxilla, Mcii, Mciii, Medial, Medial side, Median, Medifossette, Mesaceratherium, Mesaceratherium welcommi, Mesocuneiform, Mesostyle, Metacarpal, Metacone, Metaconid, Metaloph, Metapodials, Mhnt, Minutum, Miocene, Miocene rhinocerotids, Mirallesi, Mnhn, Molar, Monophyletic, Mtii, Mtiii, Mtiv, Natural history, Naturelle, Navicular, Node, Oblique, Occlusal, Occlusal view, Oligocene, Osborni, Outgroup, Paracone, Paralophid, Patella, Paulhiacense, Perissodactyla, Phylogenetic, Plesiaceratherium, Pleuroceros, Pleuroceros blanfordi, Pmnh, Postcranial, Postcranials, Posterior, Posterior border, Posterior part, Posterior tuberosity, Posterior valley, Posterolateral, Posteroproximal, Praecursor, Premolar, Processus, Processus zygomaticus, Prosantorhinus, Protaceratherium, Protocone, Protoloph, Prox, Proximal, Proximal border, Proximal view, Reversion, Rhinocerotid, Rhinocerotidae, Rhinocerotids, Rhinocerotina, Rhinocerotinae, Right astragalus, Right cuboid, Right scaphoid, Robust, Ronzotherium, Sagittal, Sagittally, Scaphoid, Semilunate, Sensu, Simorrense, Squamosal, Square brackets, Stratigraphical, Suprageneric, Symphysis, Synapomorphies, Talus, Taxon, Tibia, Transverse diameter, Trapezoid, Trigonias, Trigonid, Trochlea, Tuber, Tubercle, Tuberosity, Ulna, Unciform, Upper member, Upper molars, Upper premolars, Welcomme, Welcommi, Zinda, Zoological, Zoological journal, Zygomatic, Zygomatic arch, Zygomaticus.
Abstract
Rhinocerotids are particularly abundant and diversified in Neogene deposits of the Indian subcontinent, but their systematics is far from being well defined. Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘Aceratherium blanfordi Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: Pleuroceros blanfordi and Mesaceratherium welcommi sp. nov. Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports Pleuroceros and Mesaceratherium as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, P. blanfordi and M. welcommi are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.
Url:
DOI: 10.1111/j.1096-3642.2009.00597.x
Affiliations:
- France, Pakistan, États-Unis
- Illinois, Languedoc-Roussillon, Massachusetts, Midi-Pyrénées, Occitanie (région administrative), Île-de-France
- Cambridge (Massachusetts), Montpellier, Paris, Toulouse
- Université Harvard, Université Montpellier 2
Links toward previous steps (curation, corpus...)
- to stream Istex, to step Corpus: 001D41
- to stream Istex, to step Curation: 001D41
- to stream Istex, to step Checkpoint: 001F72
- to stream Main, to step Merge: 005511
- to stream Main, to step Curation: 005468
- to stream Main, to step Exploration: 005468
- to stream France, to step Extraction: 000134
Links to Exploration step
ISTEX:3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3Le document en format XML
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Downing</name><affiliation wicri:level="2"><country xml:lang="fr">États-Unis</country><wicri:regionArea>DePaul University, Chicago, IL 60604</wicri:regionArea><placeName><region type="state">Illinois</region></placeName></affiliation></author><author><name sortKey="Crochet, Jean Ves" sort="Crochet, Jean Ves" uniqKey="Crochet J" first="Jean-Yves" last="Crochet">Jean-Yves Crochet</name><affiliation wicri:level="4"><country xml:lang="fr">France</country><wicri:regionArea>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5</wicri:regionArea><orgName type="university">Université Montpellier 2</orgName><placeName><settlement type="city">Montpellier</settlement><region type="region" nuts="2">Occitanie (région administrative)</region><region type="old region" nuts="2">Languedoc-Roussillon</region></placeName></affiliation></author><author><name sortKey="Duranthon, Francis" sort="Duranthon, Francis" uniqKey="Duranthon F" first="Francis" last="Duranthon">Francis Duranthon</name><affiliation wicri:level="1"><country xml:lang="fr">France</country><wicri:regionArea>Laboratoire d'Étude et de Conservation du Patrimoine, Muséum d'Histoire naturelle de Toulouse, 39 allées Jules Guesde, F‐31000 Toulouse</wicri:regionArea><wicri:noRegion>31000 Toulouse</wicri:noRegion><wicri:noRegion>F‐31000 Toulouse</wicri:noRegion></affiliation></author><author><name sortKey="Flynn, Lawrence J" sort="Flynn, Lawrence J" uniqKey="Flynn L" first="Lawrence J." last="Flynn">Lawrence J. Flynn</name><affiliation wicri:level="4"><country xml:lang="fr">États-Unis</country><wicri:regionArea>Department of Anthropology and Peabody Museum, Harvard University, Cambridge, MA 02138</wicri:regionArea><placeName><region type="state">Massachusetts</region><settlement type="city">Cambridge (Massachusetts)</settlement></placeName><orgName type="university">Université Harvard</orgName></affiliation></author><author><name sortKey="Marivaux, Laurent" sort="Marivaux, Laurent" uniqKey="Marivaux L" first="Laurent" last="Marivaux">Laurent Marivaux</name><affiliation wicri:level="4"><country xml:lang="fr">France</country><wicri:regionArea>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5</wicri:regionArea><placeName><region type="region" nuts="2">Occitanie (région administrative)</region><region type="old region" nuts="2">Languedoc-Roussillon</region><settlement type="city">Montpellier</settlement><settlement type="city">Montpellier</settlement></placeName><orgName type="university">Université Montpellier 2</orgName></affiliation></author><author><name sortKey="Metais, Gregoire" sort="Metais, Gregoire" uniqKey="Metais G" first="Grégoire" last="Métais">Grégoire Métais</name><affiliation wicri:level="3"><country xml:lang="fr">France</country><wicri:regionArea>Laboratoire de Paléontologie, UMR 5143 CNRS, Muséum National d'Histoire Naturelle, 8 rue Buffon, F‐75005 Paris</wicri:regionArea><placeName><region type="region" nuts="2">Île-de-France</region><settlement type="city">Paris</settlement></placeName></affiliation></author><author><name sortKey="Rajpar, Abdul Rahim" sort="Rajpar, Abdul Rahim" uniqKey="Rajpar A" first="Abdul Rahim" last="Rajpar">Abdul Rahim Rajpar</name><affiliation wicri:level="1"><country xml:lang="fr">Pakistan</country><wicri:regionArea>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad</wicri:regionArea><wicri:noRegion>44000 Islamabad</wicri:noRegion></affiliation></author><author><name sortKey="Roohi, Ghazala" sort="Roohi, Ghazala" uniqKey="Roohi G" first="Ghazala" last="Roohi">Ghazala Roohi</name><affiliation wicri:level="1"><country xml:lang="fr">Pakistan</country><wicri:regionArea>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad</wicri:regionArea><wicri:noRegion>44000 Islamabad</wicri:noRegion></affiliation></author></analytic><monogr></monogr><series><title level="j" type="main">Zoological Journal of the Linnean Society</title><title level="j" type="alt">ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY</title><idno type="ISSN">0024-4082</idno><idno type="eISSN">1096-3642</idno><imprint><biblScope unit="vol">160</biblScope><biblScope unit="issue">1</biblScope><biblScope unit="page" from="139">139</biblScope><biblScope unit="page" to="194">194</biblScope><biblScope unit="page-count">56</biblScope><publisher>Blackwell Publishing Ltd</publisher><pubPlace>Oxford, UK</pubPlace><date type="published" when="2010-09">2010-09</date></imprint><idno type="ISSN">0024-4082</idno></series></biblStruct></sourceDesc><seriesStmt><idno type="ISSN">0024-4082</idno></seriesStmt></fileDesc><profileDesc><textClass><keywords scheme="KwdEn" xml:lang="en"><term>Aceratheriini</term><term>Aceratherium</term><term>Albigense</term><term>Alicornops</term><term>Antecrochet</term><term>Anterior</term><term>Anterior border</term><term>Anterior side</term><term>Anterior view</term><term>Anteroposterior</term><term>Anteroposterior diameter</term><term>Antoine</term><term>Antoine welcomme</term><term>Apophysis</term><term>Approximate dimensions</term><term>Articular</term><term>Articulation</term><term>Astragalus</term><term>Attened</term><term>Balochistan</term><term>Bicornis</term><term>Blanfordi</term><term>Bonis</term><term>Brachypotherium</term><term>Brachypus</term><term>Bracket</term><term>Browni</term><term>Bugti</term><term>Bugti hills</term><term>Bula</term><term>Calcaneum</term><term>Calcaneus</term><term>Carpalis</term><term>Cheek teeth</term><term>Chilotherium</term><term>Chitarwata</term><term>Cingulum</term><term>Clade</term><term>Coeval</term><term>Constricted</term><term>Convex</term><term>Corpus mandibulae</term><term>Cranial</term><term>Crista</term><term>Cuboid</term><term>Deciduous</term><term>Dentition</term><term>Dera</term><term>Dera bugti</term><term>Diaceratherium</term><term>Diaphysis</term><term>Diceratherium</term><term>Dicerorhinus</term><term>Diceros</term><term>Dihedron</term><term>Distal articulation</term><term>Distal border</term><term>Distal view</term><term>Dorsal</term><term>Dorsal view</term><term>Douvillei</term><term>Duvernoy</term><term>Earliest miocene</term><term>Early miocene</term><term>Ectocuneiform</term><term>Ectoloph</term><term>Ectometaloph</term><term>Elasmotheriina</term><term>Elasmotheriinae</term><term>Entoconid</term><term>Extensor carpalis</term><term>External groove</term><term>Facet</term><term>Fauna</term><term>Femur</term><term>Foramen</term><term>Fossil</term><term>Gaimersheimense</term><term>Gaindatherium</term><term>Gand</term><term>Gers</term><term>Ginsburg</term><term>Groove</term><term>Gutter</term><term>Heissig</term><term>Homoplastic</term><term>Hooijer</term><term>Humerus</term><term>Hypocone</term><term>Hypolophid</term><term>Incisivum</term><term>Insertion</term><term>Interossei</term><term>Kumbi</term><term>Labial</term><term>Labial cingulum</term><term>Lartet</term><term>Lateral</term><term>Lateral side</term><term>Lateral view</term><term>Laterally</term><term>Lholi</term><term>Lingual</term><term>Lingual bridge</term><term>Lingual cingulum</term><term>Lingual groove</term><term>Linnaeus</term><term>Linnean</term><term>Linnean society</term><term>Lower cheek teeth</term><term>Lower molars</term><term>Lower premolars</term><term>Lydekker</term><term>Mammal</term><term>Mammalia</term><term>Mandible</term><term>Mandibulae</term><term>Mandibular</term><term>Maxilla</term><term>Mcii</term><term>Mciii</term><term>Medial</term><term>Medial side</term><term>Median</term><term>Medifossette</term><term>Mesaceratherium</term><term>Mesaceratherium welcommi</term><term>Mesocuneiform</term><term>Mesostyle</term><term>Metacarpal</term><term>Metacone</term><term>Metaconid</term><term>Metaloph</term><term>Metapodials</term><term>Mhnt</term><term>Minutum</term><term>Miocene</term><term>Miocene rhinocerotids</term><term>Mirallesi</term><term>Mnhn</term><term>Molar</term><term>Monophyletic</term><term>Mtii</term><term>Mtiii</term><term>Mtiv</term><term>Natural history</term><term>Naturelle</term><term>Navicular</term><term>Node</term><term>Oblique</term><term>Occlusal</term><term>Occlusal view</term><term>Oligocene</term><term>Osborni</term><term>Outgroup</term><term>Paracone</term><term>Paralophid</term><term>Patella</term><term>Paulhiacense</term><term>Perissodactyla</term><term>Phylogenetic</term><term>Plesiaceratherium</term><term>Pleuroceros</term><term>Pleuroceros blanfordi</term><term>Pmnh</term><term>Postcranial</term><term>Postcranials</term><term>Posterior</term><term>Posterior border</term><term>Posterior part</term><term>Posterior tuberosity</term><term>Posterior valley</term><term>Posterolateral</term><term>Posteroproximal</term><term>Praecursor</term><term>Premolar</term><term>Processus</term><term>Processus zygomaticus</term><term>Prosantorhinus</term><term>Protaceratherium</term><term>Protocone</term><term>Protoloph</term><term>Prox</term><term>Proximal</term><term>Proximal border</term><term>Proximal view</term><term>Reversion</term><term>Rhinocerotid</term><term>Rhinocerotidae</term><term>Rhinocerotids</term><term>Rhinocerotina</term><term>Rhinocerotinae</term><term>Right astragalus</term><term>Right cuboid</term><term>Right scaphoid</term><term>Robust</term><term>Ronzotherium</term><term>Sagittal</term><term>Sagittally</term><term>Scaphoid</term><term>Semilunate</term><term>Sensu</term><term>Simorrense</term><term>Squamosal</term><term>Square brackets</term><term>Stratigraphical</term><term>Suprageneric</term><term>Symphysis</term><term>Synapomorphies</term><term>Talus</term><term>Taxon</term><term>Tibia</term><term>Transverse diameter</term><term>Trapezoid</term><term>Trigonias</term><term>Trigonid</term><term>Trochlea</term><term>Tuber</term><term>Tubercle</term><term>Tuberosity</term><term>Ulna</term><term>Unciform</term><term>Upper member</term><term>Upper molars</term><term>Upper premolars</term><term>Welcomme</term><term>Welcommi</term><term>Zinda</term><term>Zoological</term><term>Zoological journal</term><term>Zygomatic</term><term>Zygomatic arch</term><term>Zygomaticus</term></keywords><keywords scheme="Teeft" xml:lang="en"><term>Aceratheriini</term><term>Aceratherium</term><term>Albigense</term><term>Alicornops</term><term>Antecrochet</term><term>Anterior</term><term>Anterior border</term><term>Anterior side</term><term>Anterior view</term><term>Anteroposterior</term><term>Anteroposterior diameter</term><term>Antoine</term><term>Antoine welcomme</term><term>Apophysis</term><term>Approximate dimensions</term><term>Articular</term><term>Articulation</term><term>Astragalus</term><term>Attened</term><term>Balochistan</term><term>Bicornis</term><term>Blanfordi</term><term>Bonis</term><term>Brachypotherium</term><term>Brachypus</term><term>Bracket</term><term>Browni</term><term>Bugti</term><term>Bugti hills</term><term>Bula</term><term>Calcaneum</term><term>Calcaneus</term><term>Carpalis</term><term>Cheek teeth</term><term>Chilotherium</term><term>Chitarwata</term><term>Cingulum</term><term>Clade</term><term>Coeval</term><term>Constricted</term><term>Convex</term><term>Corpus mandibulae</term><term>Cranial</term><term>Crista</term><term>Cuboid</term><term>Deciduous</term><term>Dentition</term><term>Dera</term><term>Dera bugti</term><term>Diaceratherium</term><term>Diaphysis</term><term>Diceratherium</term><term>Dicerorhinus</term><term>Diceros</term><term>Dihedron</term><term>Distal articulation</term><term>Distal border</term><term>Distal view</term><term>Dorsal</term><term>Dorsal view</term><term>Douvillei</term><term>Duvernoy</term><term>Earliest miocene</term><term>Early miocene</term><term>Ectocuneiform</term><term>Ectoloph</term><term>Ectometaloph</term><term>Elasmotheriina</term><term>Elasmotheriinae</term><term>Entoconid</term><term>Extensor carpalis</term><term>External groove</term><term>Facet</term><term>Fauna</term><term>Femur</term><term>Foramen</term><term>Fossil</term><term>Gaimersheimense</term><term>Gaindatherium</term><term>Gand</term><term>Gers</term><term>Ginsburg</term><term>Groove</term><term>Gutter</term><term>Heissig</term><term>Homoplastic</term><term>Hooijer</term><term>Humerus</term><term>Hypocone</term><term>Hypolophid</term><term>Incisivum</term><term>Insertion</term><term>Interossei</term><term>Kumbi</term><term>Labial</term><term>Labial cingulum</term><term>Lartet</term><term>Lateral</term><term>Lateral side</term><term>Lateral view</term><term>Laterally</term><term>Lholi</term><term>Lingual</term><term>Lingual bridge</term><term>Lingual cingulum</term><term>Lingual groove</term><term>Linnaeus</term><term>Linnean</term><term>Linnean society</term><term>Lower cheek teeth</term><term>Lower molars</term><term>Lower premolars</term><term>Lydekker</term><term>Mammal</term><term>Mammalia</term><term>Mandible</term><term>Mandibulae</term><term>Mandibular</term><term>Maxilla</term><term>Mcii</term><term>Mciii</term><term>Medial</term><term>Medial side</term><term>Median</term><term>Medifossette</term><term>Mesaceratherium</term><term>Mesaceratherium welcommi</term><term>Mesocuneiform</term><term>Mesostyle</term><term>Metacarpal</term><term>Metacone</term><term>Metaconid</term><term>Metaloph</term><term>Metapodials</term><term>Mhnt</term><term>Minutum</term><term>Miocene</term><term>Miocene rhinocerotids</term><term>Mirallesi</term><term>Mnhn</term><term>Molar</term><term>Monophyletic</term><term>Mtii</term><term>Mtiii</term><term>Mtiv</term><term>Natural history</term><term>Naturelle</term><term>Navicular</term><term>Node</term><term>Oblique</term><term>Occlusal</term><term>Occlusal view</term><term>Oligocene</term><term>Osborni</term><term>Outgroup</term><term>Paracone</term><term>Paralophid</term><term>Patella</term><term>Paulhiacense</term><term>Perissodactyla</term><term>Phylogenetic</term><term>Plesiaceratherium</term><term>Pleuroceros</term><term>Pleuroceros blanfordi</term><term>Pmnh</term><term>Postcranial</term><term>Postcranials</term><term>Posterior</term><term>Posterior border</term><term>Posterior part</term><term>Posterior tuberosity</term><term>Posterior valley</term><term>Posterolateral</term><term>Posteroproximal</term><term>Praecursor</term><term>Premolar</term><term>Processus</term><term>Processus zygomaticus</term><term>Prosantorhinus</term><term>Protaceratherium</term><term>Protocone</term><term>Protoloph</term><term>Prox</term><term>Proximal</term><term>Proximal border</term><term>Proximal view</term><term>Reversion</term><term>Rhinocerotid</term><term>Rhinocerotidae</term><term>Rhinocerotids</term><term>Rhinocerotina</term><term>Rhinocerotinae</term><term>Right astragalus</term><term>Right cuboid</term><term>Right scaphoid</term><term>Robust</term><term>Ronzotherium</term><term>Sagittal</term><term>Sagittally</term><term>Scaphoid</term><term>Semilunate</term><term>Sensu</term><term>Simorrense</term><term>Squamosal</term><term>Square brackets</term><term>Stratigraphical</term><term>Suprageneric</term><term>Symphysis</term><term>Synapomorphies</term><term>Talus</term><term>Taxon</term><term>Tibia</term><term>Transverse diameter</term><term>Trapezoid</term><term>Trigonias</term><term>Trigonid</term><term>Trochlea</term><term>Tuber</term><term>Tubercle</term><term>Tuberosity</term><term>Ulna</term><term>Unciform</term><term>Upper member</term><term>Upper molars</term><term>Upper premolars</term><term>Welcomme</term><term>Welcommi</term><term>Zinda</term><term>Zoological</term><term>Zoological journal</term><term>Zygomatic</term><term>Zygomatic arch</term><term>Zygomaticus</term></keywords></textClass></profileDesc></teiHeader><front><div type="abstract">Rhinocerotids are particularly abundant and diversified in Neogene deposits of the Indian subcontinent, but their systematics is far from being well defined. Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘Aceratherium blanfordi Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: Pleuroceros blanfordi and Mesaceratherium welcommi sp. nov. Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports Pleuroceros and Mesaceratherium as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, P. blanfordi and M. welcommi are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.</div></front></TEI><affiliations><list><country><li>France</li><li>Pakistan</li><li>États-Unis</li></country><region><li>Illinois</li><li>Languedoc-Roussillon</li><li>Massachusetts</li><li>Midi-Pyrénées</li><li>Occitanie (région administrative)</li><li>Île-de-France</li></region><settlement><li>Cambridge (Massachusetts)</li><li>Montpellier</li><li>Paris</li><li>Toulouse</li></settlement><orgName><li>Université Harvard</li><li>Université Montpellier 2</li></orgName></list><tree><country name="France"><region name="Occitanie (région administrative)"><name sortKey="Antoine, Pierre Livier" sort="Antoine, Pierre Livier" uniqKey="Antoine P" first="Pierre-Olivier" last="Antoine">Pierre-Olivier Antoine</name></region><name sortKey="Antoine, Pierre Livier" sort="Antoine, Pierre Livier" uniqKey="Antoine P" first="Pierre-Olivier" last="Antoine">Pierre-Olivier Antoine</name><name sortKey="Antoine, Pierre Livier" sort="Antoine, Pierre Livier" uniqKey="Antoine P" first="Pierre-Olivier" last="Antoine">Pierre-Olivier Antoine</name><name sortKey="Antoine, Pierre Livier" sort="Antoine, Pierre Livier" uniqKey="Antoine P" first="Pierre-Olivier" last="Antoine">Pierre-Olivier Antoine</name><name sortKey="Crochet, Jean Ves" sort="Crochet, Jean Ves" uniqKey="Crochet J" first="Jean-Yves" last="Crochet">Jean-Yves Crochet</name><name sortKey="Duranthon, Francis" sort="Duranthon, Francis" uniqKey="Duranthon F" first="Francis" last="Duranthon">Francis Duranthon</name><name sortKey="Marivaux, Laurent" sort="Marivaux, Laurent" uniqKey="Marivaux L" first="Laurent" last="Marivaux">Laurent Marivaux</name><name sortKey="Metais, Gregoire" sort="Metais, Gregoire" uniqKey="Metais G" first="Grégoire" last="Métais">Grégoire Métais</name></country><country name="États-Unis"><region name="Illinois"><name sortKey="Downing, Kevin F" sort="Downing, Kevin F" uniqKey="Downing K" first="Kevin F." last="Downing">Kevin F. Downing</name></region><name sortKey="Flynn, Lawrence J" sort="Flynn, Lawrence J" uniqKey="Flynn L" first="Lawrence J." last="Flynn">Lawrence J. Flynn</name></country><country name="Pakistan"><noRegion><name sortKey="Rajpar, Abdul Rahim" sort="Rajpar, Abdul Rahim" uniqKey="Rajpar A" first="Abdul Rahim" last="Rajpar">Abdul Rahim Rajpar</name></noRegion><name sortKey="Roohi, Ghazala" sort="Roohi, Ghazala" uniqKey="Roohi G" first="Ghazala" last="Roohi">Ghazala Roohi</name></country></tree></affiliations></record>Pour manipuler ce document sous Unix (Dilib)
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